Virkki, N., and J. A. Santiago-Blay. “Chromosome Numbers in 71 Puerto Rican Species of Leaf Beetles (Coleoptera: Chrysomelidae)”. The Journal of Agriculture of the University of Puerto Rico, vol. 82, no. 1-2, Apr. 1998, pp. 69-83, doi:10.46429/jaupr.v82i1-2.3821.